Mike Elliott will replace James Allison as technical director in a reshuffle for Formula One constructors' champions Mercedes.
Allison has led the Silver Arrows' technical operations since early in 2017, but will step back from day-to-day management to become chief technical officer on July 1.
Elliott, who started his career with McLaren 21 years ago and also worked for Renault, will be promoted to replace Allison.
He joined Mercedes as head of aerodynamics in 2012 before being appointed as the team's technology director four years ago and will switch jobs again in another reshuffle.
Mercedes said in a statement on Friday: "The transition to the new organisation will take place across the coming months as we continue the 2021 championship battle on track and prepare for the significant challenge of the 2022 technical and sporting regulations in our development programmes back at base."
Elliott said: "It has been a great pleasure and privilege to work for James earlier in my career at Renault, and for the past four years at Mercedes. His track record in the sport speaks for itself, and he has been a fantastic team-mate and leader for me during that time.
"They are big shoes to fill and I am delighted that we will be able to call on his expertise in his new role as CTO. On a personal level, it is an amazing opportunity to become Technical Director of a team like Mercedes, and I must thank Toto [Wolff] and our company for trusting me to make the step up to this new role.
"It is an incredible privilege to be part of this team and I know that the leadership strength at every level through the company will be vital to our future success.
"I can't wait to get started and to tackle the many exciting technical challenges ahead of us in the next months and years."
The Silver Arrows have won seven consecutive constructors' titles and seven-time world champion Lewis Hamilton won the season-opening Bahrain Grand Prix last month.