Sean O'Loughlin will not be included in the Great Britain Lions' tour of New Zealand and Papua New Guinea, coach Wayne Bennett has confirmed.

O'Loughlin was named in Bennett's most recent training squad ahead of the four-match tour, which starts against Tonga in Hamilton on October 26.

However, the England and Wigan Warriors captain - who played in the last Great Britain side in 2007 - will not feature in the travelling party after a pectoral injury restricted him to 18 Super League appearances this term.

Tom Makinson joins the 36-year-old in sitting out the tour, having dislocated his shoulder in St Helens' Grand Final win over Salford at Old Trafford on Saturday.

"I've admired Sean as a player for a long time, and I've thoroughly enjoyed working with him with England," Bennett said of O'Louglin.

"He's been an outstanding leader of the team on and off the field, with the ability to play in any position, and his combination of skill and toughness.

"It was important that I spoke to him and we agreed on an outcome, and hopefully it will help him come back fresh next season."

James Graham seems the likely candidate to replace O'Loughlin as captain, with Sam Burgess having also pulled out of the squad due to a shoulder injury.